{"id":233677,"date":"2025-09-16T11:44:35","date_gmt":"2025-09-16T10:44:35","guid":{"rendered":"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/?post_type=vf_event&#038;p=233677"},"modified":"2026-08-19T15:37:25","modified_gmt":"2026-08-19T14:37:25","slug":"ees27-03","status":"publish","type":"vf_event","link":"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/events\/ees27-03\/","title":{"rendered":"Evolutionary biochemistry and molecular biology: origin and diversification of molecular systems"},"content":{"rendered":"\n<div class=\"vf-tabs\"><ul class=\"vf-tabs__list\" data-vf-js-tabs=\"true\"><li class=\"vf-tabs__item\"><a class=\"vf-tabs__link\" href=\"#vf-tabs__section-overview\" data-vf-js-location-nearest-activation-target=\"\">Overview<\/a><\/li><li class=\"vf-tabs__item\"><a class=\"vf-tabs__link\" href=\"#vf-tabs__section-921bb823-d78a-49dc-a09e-4fbd29733cba\" data-vf-js-location-nearest-activation-target=\"\">Speakers<\/a><\/li><li class=\"vf-tabs__item\"><a class=\"vf-tabs__link\" href=\"#vf-tabs__section-sponsors\" data-vf-js-location-nearest-activation-target=\"\">Sponsors<\/a><\/li><li class=\"vf-tabs__item\"><a class=\"vf-tabs__link\" href=\"#vf-tabs__section-about\" data-vf-js-location-nearest-activation-target=\"\">About<\/a><\/li><li class=\"vf-tabs__item\"><a class=\"vf-tabs__link\" href=\"#vf-tabs__section-mediakit\" data-vf-js-location-nearest-activation-target=\"\">Media kit<\/a><\/li><\/ul><div class=\"vf-tabs-content\" data-vf-js-tabs-content=\"true\">\n<section class=\"vf-tabs__section\" id=\"vf-tabs__section-overview\"><h2 class=\"vf-u-sr-only\">Overview<\/h2>\n<p class=\"wp-block-paragraph\">This conference will take place at EMBL Heidelberg, with the option to attend virtually. <\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Registration is not yet open for this event. If you are interested in receiving more information please register your interest. <\/p>\n\n\n\n<h2 class=\"wp-block-heading\">Symposium overview<\/h2>\n\n\n\n<p class=\"wp-block-paragraph\">Life&#8217;s history is written in molecules. This symposium will explore how the molecular systems of modern cells originated, evolved, and gave rise to biological complexity.<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Evolutionary biochemistry is emerging as a transformative field. Advances in genomics, AI-enabled protein analysis, structural biology, and ancestral sequence reconstruction now allow researchers to investigate the origins of proteins, metabolism, and cells with unprecedented precision, making this an ideal moment to unite the field.<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">By bringing together researchers in chemistry, biochemistry, evolutionary biology, and computational science, this meeting will foster collaborations, establish shared approaches, and help define the future directions of evolutionary biochemistry.<\/p>\n\n\n\n<h2 class=\"wp-block-heading\">Session topics<\/h2>\n\n\n\n<ul class=\"wp-block-list\">\n<li>Origins of biological chemistry and early metabolism<\/li>\n\n\n\n<li>Evolution of protein structure and function<\/li>\n\n\n\n<li>Evolution of metabolic networks<\/li>\n\n\n\n<li>Evolutionary cell biology<\/li>\n\n\n\n<li>Emergence of molecular complexity<\/li>\n\n\n\n<li>Computational and experimental methods in evolutionary biochemistry<\/li>\n\n\n\n<li>Ancestral sequence reconstruction and protein resurrection<\/li>\n\n\n\n<li>AI, structural biology, and comparative genomics for evolutionary discovery<\/li>\n<\/ul>\n\n\n\n<h2 class=\"wp-block-heading\">Who should attend<\/h2>\n\n\n\n<p class=\"wp-block-paragraph\">Designed for researchers at all career stages with an interest in molecular evolution and the origins of biological systems.<\/p>\n\n\n\n<p class=\"wp-block-paragraph\"><em>EMBL Events is committed to creating an inclusive environment where internationally renowned researchers and researchers at all career stages across the life sciences share knowledge, build connections, and foster collaboration.<\/em><\/p>\n<\/section>\n\n\n\n<section class=\"vf-tabs__section\" id=\"vf-tabs__section-921bb823-d78a-49dc-a09e-4fbd29733cba\"><h2>Speakers<\/h2>\n<div class=\"embl-grid\"><div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"300\" height=\"300\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Baum_Buzz-1.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Baum_Buzz-1.jpg 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Baum_Buzz-1-150x150.jpg 150w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Buzz\tBaum            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                MRC Laboratory of Molecular Biology            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                UK            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"225\" height=\"300\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Beltrao_Pedro-225x300.jpeg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Beltrao_Pedro-225x300.jpeg 225w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Beltrao_Pedro.jpeg 300w\" sizes=\"auto, (max-width: 225px) 100vw, 225px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Pedro\tBeltrao            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                ETH Z\u00fcrich            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Switzerland            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"300\" height=\"211\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Carvunis_Anne-e1760017701486-300x211.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Carvunis_Anne-e1760017701486-300x211.jpg 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Carvunis_Anne-e1760017701486.jpg 413w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Anne-Ruxandra\tCarvunis            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                University of Pittsburgh            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                USA            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n<\/div>\n\n\n\n<div class=\"embl-grid\"><div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"300\" height=\"300\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Ferreiro-Diego.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Ferreiro-Diego.jpg 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Ferreiro-Diego-150x150.jpg 150w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Diego U.\tFerreiro            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                University of Buenos Aires            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Argentina            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"225\" height=\"225\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Hlouchova_Klara.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Hlouchova_Klara.jpg 225w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Hlouchova_Klara-150x150.jpg 150w\" sizes=\"auto, (max-width: 225px) 100vw, 225px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Kl\u00e1ra\tHlouchov\u00e1            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Charles University            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Czech Republic            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"120\" height=\"150\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Kacar_Betul.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Bet\u00fcl Ka\u00e7ar            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                University of Wisconsin\u2013Madison            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                USA            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n<\/div>\n\n\n\n<div class=\"embl-grid\"><div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"194\" height=\"146\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Kamerlin_Lynn-e1760017738689.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Lynn\tKamerlin            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Georgia Institute of Technology            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                USA            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"300\" height=\"290\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Kurth_Julia-300x290.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Kurth_Julia-300x290.jpg 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Kurth_Julia.jpg 430w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Julia\tKurth            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                University of M\u00fcnster            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Germany            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"183\" height=\"275\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Levy_Emmanuel.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Emmanuel\tLevy            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                University of Geneva            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Switzerland            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n<\/div>\n\n\n\n<div class=\"embl-grid\"><div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"233\" height=\"256\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Longo-Lian.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Liam\tLongo            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Earth-Life Science Institute, Institute of Science Tokyo            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Japan            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"300\" height=\"300\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Moran_Joseph-300x300.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Moran_Joseph-300x300.jpg 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Moran_Joseph-150x150.jpg 150w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Moran_Joseph-768x768.jpg 768w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Moran_Joseph.jpg 930w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Joseph\tMoran            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                University of Ottawa            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Canada            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"180\" height=\"280\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Munoz-Gomez_Sergio.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Sergio\tMu\u00f1oz-G\u00f3mez            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Purdue University            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                USA            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n<\/div>\n\n\n\n<div class=\"embl-grid\"><div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"300\" height=\"300\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Noda_Lianet-300x300.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Noda_Lianet-300x300.jpg 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Noda_Lianet-150x150.jpg 150w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Noda_Lianet-768x768.jpg 768w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Noda_Lianet.jpg 1024w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Lianet\tNoda Garcia            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                The Hebrew University of Jerusalem            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Israel            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"225\" height=\"225\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Pinney_Margaux.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Pinney_Margaux.jpg 225w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Pinney_Margaux-150x150.jpg 150w\" sizes=\"auto, (max-width: 225px) 100vw, 225px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Margaux M.\tPinney            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                University of California, Berkeley            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                USA            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"225\" height=\"225\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Steinegger.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Steinegger.jpg 225w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Steinegger-150x150.jpg 150w\" sizes=\"auto, (max-width: 225px) 100vw, 225px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Martin\tSteinegger            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Seoul National University            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                South Korea            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n<\/div>\n\n\n\n<div class=\"embl-grid\"><div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"300\" height=\"300\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Thornton_Josepf-300x300.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Thornton_Josepf-300x300.jpg 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Thornton_Josepf-150x150.jpg 150w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Thornton_Josepf.jpg 600w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Joseph\tThornton            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                The University of Chicago            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                USA            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"300\" height=\"300\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Tokuriki-Nobuhiko-300x300.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Tokuriki-Nobuhiko-300x300.jpg 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Tokuriki-Nobuhiko-150x150.jpg 150w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Tokuriki-Nobuhiko.jpg 411w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Nobuhiko\tTokuriki            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                The University of British Columbia            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Canada            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"300\" height=\"300\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Toth-Petroczy_Agnes-1-300x300.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Toth-Petroczy_Agnes-1-300x300.jpg 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Toth-Petroczy_Agnes-1-150x150.jpg 150w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Toth-Petroczy_Agnes-1.jpg 600w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Agnes\tToth-Petroczy            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Max Planck Institute of Molecular Cell Biology and Genetics            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Germany            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n<\/div>\n\n\n\n<div class=\"embl-grid\"><div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"247\" height=\"204\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Vila_Alejandro.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Alejandro\tVila            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                IBR \u2013 Instituto de Biolog\u00eda Molecular y Celular de Rosario            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Argentina            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<\/div>\n<\/div>\n<\/div>\n\n\n\n<h2 class=\"wp-block-heading\">Scientific organisers<\/h2>\n\n\n\n<div class=\"embl-grid\"><div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"155\" height=\"175\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Gautam-Dey.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Gautam\tDey            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                EMBL Heidelberg            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Germany            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"300\" height=\"232\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Hochberg_Gerog-300x232.webp\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Hochberg_Gerog-300x232.webp 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Hochberg_Gerog.webp 456w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Georg\tHochberg            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Philipps-Universit\u00e4t Marburg            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Germany            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"300\" height=\"278\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/MAL2064-scaled-e1787149853417-300x278.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/MAL2064-scaled-e1787149853417-300x278.jpg 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/MAL2064-scaled-e1787149853417-1024x950.jpg 1024w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/MAL2064-scaled-e1787149853417-768x713.jpg 768w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/MAL2064-scaled-e1787149853417.jpg 1290w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Laura\tMascotti            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Instituto de Histolog\u00eda y Embriolog\u00eda de Mendoza            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Argentina            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n<\/div>\n\n\n\n<div class=\"embl-grid\"><div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<article class=\"vf-profile vf-profile--very-easy vf-profile--medium vf-profile--block | vf-u-margin__bottom--400\">\n\n    <img decoding=\"async\" width=\"264\" height=\"300\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Markus-Ralser-264x300.jpg\" class=\"vf-profile__image\" alt=\"\" loading=\"lazy\" itemprop=\"image\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Markus-Ralser-264x300.jpg 264w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/Markus-Ralser.jpg 297w\" sizes=\"auto, (max-width: 264px) 100vw, 264px\" \/>\n    <h3 class=\"vf-profile__title\" style=\"text-align: center;\">\n                    Markus\tRalser            <\/h3>\n    \n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Charit\u00e9 Universit\u00e4tsmedizin Berlin            <\/p>\n                <p class=\"vf-profile__text\" style=\"text-align: center;\">\n                Germany            <\/p>\n    \n    \n      \n\n    \n<\/article>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<\/div>\n<\/div>\n<\/div>\n<\/section>\n\n\n\n<section class=\"vf-tabs__section\" id=\"vf-tabs__section-sponsors\"><h2>Sponsors<\/h2>\n<p class=\"wp-block-paragraph\"><\/p>\n\n\n\n<p class=\"wp-block-paragraph\"><strong>Sponsorship opportunities<\/strong><\/p>\n\n\n\n<p class=\"wp-block-paragraph\">We offer a variety of event sponsoring possibilities, with the flexibility to select a set sponsorship package or combine individual sponsorship options to suit your event budget.&nbsp;Discounts are available for companies sponsoring multiple events at EMBL Heidelberg.&nbsp;<a href=\"http:\/\/www.embl.de\/training\/events\/index.php?p_eventType=CON&amp;amp;p_outstation=HD&amp;amp;p_timeRange=FUTURE&amp;amp;submit=Show+Events\">View other conferences<\/a>, or contact&nbsp;<a href=\"mailto:sponsorship@embl.de\">sponsorship@embl.de<\/a>&nbsp;for further information.<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">If you are interested in becoming a media partner of this event, please visit our&nbsp;<a href=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/sponsor-an-event\/#vf-tabs__section-08208670-4ad4-45ca-a4ff-ac6edb6ffd0b\" target=\"_blank\" rel=\"noreferrer noopener\">media partnerships webpage<\/a>.<\/p>\n\n\n\n<div class=\"wp-block-buttons is-layout-flex wp-block-buttons-is-layout-flex\"><p><a href=\"mailto:sponsorship@embl-em.de?subject=%20Request%20a%20Sponsorship%20Brochure&#038;body=Please%20kindly%20send%20me%20the%20sponsorship%20brochure%20for%20(please%20indicate%20the%20event%20your%20are%20interested%20in).\" class=\"vf-button vf-button--primary vf-button--pill\">Request a sponsorship brochure<\/a><\/p><\/div>\n\n\n\n<div class=\"vf-box vf-box--normal vf-box-theme--quaternary | vf-u-margin__bottom--400\">\n      <h3 class=\"vf-box__heading\">\n                Warning                  <\/h3> \n        <p class=\"vf-box__text\">EMBL advises sponsors to beware of fraudulent schemes falsely offering sponsorship opportunities. All legitimate communication about sponsorship for EMBL conferences, symposia, and courses comes directly from EMBL via official EMBL email accounts. EMBL does not use third parties for sponsorship acquisition.<\/p>\n<p class=\"vf-box__text\">\u2013 EMBL does not sell attendee lists.<br \/>\n\u2013 EMBL will never ask for credit card details or direct payments.<br \/>\n\u2013 All payments are processed by invoice only.<\/p>\n<p class=\"vf-box__text\">If you receive a suspicious message claiming to be from or on behalf of EMBL, please report it to\u00a0<a href=\"mailto:sponsorship@embl.org\">sponsorship@embl.org<\/a>.<\/p>\n<\/div>\n<\/section>\n\n\n\n<section class=\"vf-tabs__section\" id=\"vf-tabs__section-about\"><h2>About<\/h2>\n<p class=\"wp-block-paragraph\">EMBO | EMBL Symposia promote scientific communication and collaboration in the European research area. They provide scientists with a platform to discuss and exchange ideas on forward-looking topics and new developments in the life sciences.<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">Topics emphasise upcoming developments and the interdisciplinary nature of related fields. Jointly funded and organised by&nbsp;<a href=\"https:\/\/www.embo.org\/\">EMBO&nbsp;<\/a>and&nbsp;<a href=\"https:\/\/www.embl.org\/\">EMBL&nbsp;<\/a>\u2013 and complementary to their respective courses, workshops, and conference programmes \u2013 the symposia promote scientific communication and collaboration.<\/p>\n\n\n\n<p class=\"wp-block-paragraph\">All symposia are held in the&nbsp;<a href=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/embl-advanced-training-centre-atc\/\">EMBL Advanced Training Centre<\/a>&nbsp;(ATC) in Heidelberg, Germany, or virtually.<\/p>\n<\/section>\n\n\n\n<section class=\"vf-tabs__section\" id=\"vf-tabs__section-mediakit\"><h2>Media kit<\/h2>\n<p class=\"wp-block-paragraph\">Want to let others know you&#8217;re attending this event? Take a look at our shareable media and feel free to use them in your social media channels or presentations.<\/p>\n\n\n\n<div class=\"vf-grid | vf-grid__col-3\"><div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<figure class=\"vf-figure wp-block-image  | vf-figure--align vf-figure--align-centered  size-medium is-resized\"><a href=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Story.jpg\"><img loading=\"lazy\" decoding=\"async\" width=\"169\" height=\"300\" class=\"vf-figure__image\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Story-169x300.jpg\" alt=\"\" class=\"wp-image-282301\" style=\"width:95px;height:160px\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Story-169x300.jpg 169w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Story-576x1024.jpg 576w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Story-768x1365.jpg 768w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Story-864x1536.jpg 864w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Story.jpg 1080w\" sizes=\"auto, (max-width: 169px) 100vw, 169px\" \/><\/a><figcaption class=\"vf-figure__caption\">Social 1080&#215;1920<\/figcaption><\/figure>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<figure class=\"vf-figure wp-block-image size-large is-resized is-style-default\"><img loading=\"lazy\" decoding=\"async\" width=\"819\" height=\"1024\" class=\"vf-figure__image\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Media-Kit-819x1024.jpg\" alt=\"\" class=\"wp-image-282305\" style=\"width:256px\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Media-Kit-819x1024.jpg 819w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Media-Kit-240x300.jpg 240w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Media-Kit-768x960.jpg 768w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-IG-Media-Kit.jpg 1080w\" sizes=\"auto, (max-width: 819px) 100vw, 819px\" \/><figcaption class=\"vf-figure__caption\">Social 1080&#215;1350<\/figcaption><\/figure>\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<figure class=\"vf-figure wp-block-image size-medium is-resized\"><a href=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-FB.jpg\"><img loading=\"lazy\" decoding=\"async\" width=\"300\" height=\"169\" class=\"vf-figure__image\" src=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-FB-300x169.jpg\" alt=\"\" class=\"wp-image-282309\" style=\"width:300px;height:169px\" srcset=\"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-FB-300x169.jpg 300w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-FB-1024x576.jpg 1024w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-FB-768x432.jpg 768w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-FB-1536x864.jpg 1536w, https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-FB.jpg 1600w\" sizes=\"auto, (max-width: 300px) 100vw, 300px\" \/><\/a><figcaption class=\"vf-figure__caption\">Social 1600&#215;900<\/figcaption><\/figure>\n\n<\/div>\n<\/div>\n<\/div>\n\n\n\n<div class=\"vf-grid | vf-grid__col-3\"><div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<\/div>\n<\/div>\n\n\n<div><!--[vf\/content]-->\n<div class=\"vf-content\">\n\n<\/div>\n<\/div>\n<\/div>\n<\/section>\n<\/div><\/div>\n\n\n\n<p class=\"wp-block-paragraph\"><\/p>\n","protected":false},"excerpt":{"rendered":"","protected":false},"parent":0,"menu_order":0,"template":"","embl_taxonomy":[],"class_list":["post-233677","vf_event","type-vf_event","status-publish","hentry"],"acf":[],"embl_taxonomy_terms":[],"vf_event_start_date":{"value":"20270629","formatted":"29 June 2027"},"vf_event_end_date":{"value":"20270702","formatted":"2 July 2027"},"vf_event_submission_opening":{"value":"","formatted":""},"vf_event_submission_closing":{"value":"","formatted":""},"vf_event_registration_opening":{"value":"","formatted":""},"vf_event_registration_closing":{"value":"","formatted":""},"vf_event_summary":{"value":"","formatted":""},"vf_event_additional_info":{"value":"","formatted":""},"vf_event_registration_fee":{"value":"","formatted":""},"vf_event_canceled":{"value":"no","formatted":"no"},"vf_event_venue":{"value":"EMBL Advanced Training Centre","formatted":"EMBL Advanced Training 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Georg Hochberg, Laura Mascotti, Markus Ralser","formatted":"Gautam Dey, Georg Hochberg, Laura Mascotti, Markus Ralser"},"vf_event_displayed":{"value":"EMBO | EMBL Symposium","formatted":"EMBO | EMBL Symposium"},"vf_event_subjects_training":{"value":["dbe","sb","cbb"],"formatted":[{"value":"dbe","label":"Developmental Biology and Evolution"},{"value":"sb","label":"Structural Biology"},{"value":"cbb","label":"Cell Biology and 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13:55:17","menu_order":0,"mime_type":"image\/jpeg","type":"image","subtype":"jpeg","icon":"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-includes\/images\/media\/default.png","width":2083,"height":2083,"sizes":{"thumbnail":"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-150x150.jpg","thumbnail-width":150,"thumbnail-height":150,"medium":"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-300x300.jpg","medium-width":300,"medium-height":300,"medium_large":"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-768x768.jpg","medium_large-width":768,"medium_large-height":768,"large":"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-1024x1024.jpg","large-width":1024,"large-height":1024,"1536x1536":"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-1536x1536.jpg","1536x1536-width":1536,"1536x1536-height":1536,"2048x2048":"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03-2048x2048.jpg","2048x2048-width":2048,"2048x2048-height":2048,"vf-hero":"https:\/\/www.embl.org\/about\/info\/course-and-conference-office\/wp-content\/uploads\/EES27-03.jpg","vf-hero-width":2083,"vf-hero-height":2083}}},"vf_event_custom_overview":{"value":"This symposium will explore how the molecular systems of modern cells originated, evolved, and gave rise to biological complexity. 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